双管藻属Amphisolenia与华鳍藻属Sinophysis的一些物种内共生了蓝藻。论文中通常把他们描述为盗质型内共生。呃，我觉得不算（因为蓝藻本身就是一个质体，而不是消化的只剩质体），尽管他们内共生同样不长久。

那么，怎样才能长久呢？
当然是像最初的吸噬虫祖先一样，吞噬型内共生咯
多甲藻目的克里藻科Kryptoperidiniaceae吞噬并内共生了硅藻，他们的内共生是永久的
他们的质体是5层囊膜的，硅藻的细胞膜一层再加上硅藻质体的四层（淡色藻门Ochrophyta都就具有四层囊膜的质体）（淡色藻门也属于SAR超系群生物，它们能光合同样是内共生着红藻）
由于他们内共生事件并没发生太久，所以还保留了硅藻的细胞核

裸甲藻目的鳞甲藻属Lepidodinium内共生了绿藻（属于植物），他们的质体也是永久的。鳞甲藻仍保留了绿藻的细胞核

短甲藻目的凯伦藻科Kareniaceae内共生了定鞭藻［26］，他的内共生也是永久的。与前两者不同，他并没有留下定鞭藻的细胞核
（定鞭藻门Haptophyta，也不是植物，而是属于定鞭界Haptista）
诶，桨兰你上面不是说凯伦藻科的是盗质体吗？怎么到这又成吞噬内共生了？
，不是，那个未知种确实是盗质体，但是其他凯伦藻的确实是吞噬共生——他们拥有四层囊膜，一层定鞭藻细胞膜，再加上定鞭藻门质体的三层囊膜。
两者是不同的尝试，那个未知种是盗定鞭藻质体，而其他则是由吞噬内共生定鞭藻的祖先演化而来的
顺便说一下，定鞭藻本身也是盗质体内共生的［27］，现生定鞭藻均由数亿年前一个盗隐藻质体的祖先演化而来的（图3）。好吧好吧，兜兜转转，终是用隐藻的质体
（那怎么不说终是用红藻的质体呢…不对，终是内共生蓝细菌）
一些凯伦藻科的，比如说短凯伦藻Karenia brevis会产生神经性贝毒，中毒症状为口唇、舌、咽喉刺痛，随后出现四肢麻木、头晕、恶心、呕吐、腹痛、腹泻，严重者会呼吸麻痹、昏迷甚至死亡
图4为壮观的凯伦藻赤潮（一片地狱风光

那么有没有不长久的吞噬内共生呢？当然有啊
比如说夜光藻Noctiluca scintillans就可以通过内共生梨甲藻获得光合能力
几大光合类群，植物，淡色藻，蓝细菌，定鞭藻，隐藻全都要内共生一遍，甚至是自己吸噬虫共门都不放过吗？哈基甲，你这家伙（甚至是发光甲藻内共生发光甲藻）

结语
吸噬虫亚门为何会这么多次质体丢失光合能力？这么多次演化成寄生虫？难道他们真是如此的邪恶？
不，更有可能的是，那些基干的光合吸噬虫都因无法与后来出现的更加强大的海洋光合藻类竞争而灭绝。只有那些演化为寄生虫的活了下来。因此造成了基干的吸噬虫全都演化为了寄生虫的“假像”
再说了，即使丢失了光合能力，捕食性甲藻照样有机会通过内共生再次获得光合能力
生物演化本就在不断的变化之中，唯有适者才能生存

致歉：本文章写了长达三周，是抽空一有时间就写点。导致文章不流畅，很多地方矛盾生硬，我脑子也有点问题。很抱歉的是，本期文章单纯只从分子角度来看单减虫的演化，根本没有顾及化石（没时间理了）。

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